332. Rettelbach, A., Nater, A. & Ellegren, H. (2019) How linked selection shapes the diversity landscape in Ficedula flycatchers. Genetics, in press.

331. Smeds, L., Kojola, I. & Ellegren, H. (2019) The evolutionary history of grey wolf Y chromosomes. Molecular Ecology, in press.

330. Nadachowska-Brzyska, K., Burri, R. & Ellegren, H. (2019) Footprints of adaptive evolution revealed by whole Z chromosome haplotypes in flycatchers. Molecular Ecology, in press.

329. Warmuth, V.M. & Ellegren, H. (2019) Genotype-free estimation of allele frequencies reduces bias and improves demographic inferences from RADSeq data. Molecular Ecology Resources, in press.

328. Bolívar, P., Guéguen, L., Duret, L., Ellegren, H. & Mugal, C.F. (2019) GC-biased gene conversion conceals the prediction of the nearly neutral theory in avian genomes. Genome Biology 20:5. 


327. Bolívar, P., Mugal, C.F., Rossi, M., Nater, A., Wang, M., Dutoit, L. & Ellegren, H. (2018) Biased inference of selection due to GC-biased gene conversion and the rate of protein evolution in flycatchers when accounting for it. Molecular Biology and Evolution 35:2475-2486.

326. Yazdi, H.P. & Ellegren, H. (2018) A genetic map of the ostrich Z chromosome and the role of inversions in avian sex chromosome evolution. Genome Biology and Evolution 10:2049-2060.

325. Dutoit, L., Mugal, C.F., Bolivar, P., Wang, M., Nadachowska-Brzyska, K., Smeds, L., Papoli, H., Gustafsson, L. & Ellegren, H. (2018) Sex-biased gene expression, sexual antagonism and levels of genetic diversity in the collared flycatcher (Ficedula albicollis) genome. Molecular Ecology 27:3572-3581.

324. Ekblom, R., Brechlin, B., Persson, J., Smeds, L., Johansson, M., Magnusson, J., Flagstad, Ø. & Ellegren, H. (2018). Genome sequencing and conservation genomics in the Scandinavian wolverine (Gulo gulo) population. Conservation Biology 32:1301-1312.

323. Craig, R., Suh, A., Wang, M. & Ellegren, H. (2018) Natural selection beyond genes: Identification and analyses of evolutionarily conserved elements in the genome of the collared flycatcher (Ficedula albicollis). Molecular Ecology 27:476-492.

322. Suh, A., Smeds, L. & Ellegren, H. (2018) Abundant recent activity of retrovirus-like retrotransposons within and among flycatcher species implies a rich source of structural variation in songbird genomes. Molecular Ecology 27:99-111.

321. Kardos, M., Åkesson, M., Fountain, T., Flagstad, Ø., Liberg, O., Olason, P., Sand, H., Wabakken, P., Wikenros, C. & Ellegren, H. (2018) Genomic consequences of intensive inbreeding in an isolated wolf population. Nature Ecology & Evolution 2:124-131.


320. Kawakami, T., Mugal, C.M., Suh, A., Nater, A., Burri, R., Smeds, L. & Ellegren, H. (2017) Whole-genome patterns of linkage disequilibrium across flycatcher populations clarify the causes and consequences of fine-scale recombination rate variation in birds. Molecular Ecology 26: 4158-4172.

319. Ellegren, H. & Wolf, J.B (2017) Parallelism in genomic landscapes of differentiation, conserved genomic features and the role of linked selection. Journal of Evolutionary Biology 30: 1516-1518.

318. Vijay, N., Weissensteiner, M., Burri, R., Kawakami, T., Ellegren, H. & Wolf, J.B.W. (2017) Genomewide patterns of variation in genetic diversity are shared among populations, species and higher-order taxa. Molecular Ecology 26: 4284-4295.

317. Wang, M., Uebbing, S., & Ellegren, H. (2017) Bayesian inference of allele-specific gene expression indicates abundant cis-regulatory variation in natural flycatcher populations. Genome Biology and Evolution 9: 1266-1279.

316. Silva, C.N.S, McFarlane, S.E., Hagen, I.J., Rönnegård, L., Billing, A.M., Kvalnes, T., Kemppainen, P., Rønning, B., Ringsby, T.H., Sæther, B-E., Qvarnström, A., Ellegren, H., Jensen, H, & Husby, A. (2017) Insights into the genetic architecture of morphological traits in two passerine bird species. Heredity 119: 197-205.

315. Dutoit, L., Vijay, N., Mugal, C.F., Bossu, C.M., Burri, R., Wolf, J. & Ellegren, H. (2017) Co-variation in levels of nucleotide diversity in homologous regions of the avian genome long after completion of lineage sorting. Proceedings of the Royal Society of London Series B: Biological Sciences 284: 20162756.

314. Kardos, M., Qvarnström, A., & Ellegren, H. (2017) Inferring individual inbreeding and demographic history from segments of identity by descent in Ficedula flycatcher genome sequences. Genetics 205: 1319-1334.

313. Knief, U., Schielzeth, H., Backström, N., Hemmrich-Stanisak, G., Wittig, M., Franke, A., Griffith, S.C., Ellegren, H., Kempenaers, B. & Forstmeier, W. (2017) Association mapping of morphological traits in wild and captive zebra finches: reliable within but not between populations. Molecular Ecology 26: 1285-1305.

312. Dutoit, L. Burri, R., Nater, A., Mugal, C.F. & Ellegren, H. (2017) Genomic distribution and estimation of nucleotide diversity in natural populations: perspectives from the collared flycatcher (Ficedula albicollis) genome. Molecular Ecology Resources 17:586-597.

311. Wolf, J.B. & Ellegren, H. (2017) Making sense of genomic islands of differentiation in light of speciation. Nature Reviews Genetics  18:87-100.


310. Kardos, M., Taylor, H.R., Ellegren, H., Luikart, G. & Allendorf, F.W. (2016) Genomics advances the study of inbreeding depression in the wild. Evolutionary Applications 9:1205-1218.

309. Smeds, L., Qvarnström, A. & Ellegren, H. (2016) Direct estimate of the rate of germline mutation in a bird. Genome Research 26:1211-128.

308. Smeds, L., Mugal, C.F., Qvarnström, A. & Ellegren, H. (2016) High-resolution mapping of crossover and non-crossover recombination events by whole-genome re-sequencing of an avian pedigree. PLoS Genetics 12:e1006044.

307. Ellegren, H. & Galtier, N. (2016) Determinants of genetic diversity. Nature Reviews Genetics 17:422-433.

306. Uebbing, S., Künstner, A., Mäkinen, H., Backström, N., Bolivar, P., Burri, R., Dutoit, L., Mugal, C.F., Nater, A., Aken, B., Flicek, P., Martin, F.J., Searle, S.M.J. & Ellegren, H. (2016) Divergence in gene expression within and between two closely related flycatcher species. Molecular Ecology 25:2015-2028.

305. Figuet, E., Nabholz, B., Bonneau, M., Mas Carrio, E., Nadachowska-Brzyska, K., Ellegren, H. & Galiter, N. (2016) Life history traits, protein evolution and the nearly-neutral theory in amniotes. Molecular Biology and Evolution 33:1517-1527.

304. Nadachowska-Brzyska, K., Burri, R., Smeds, L. & Ellegren, H. (2016) PSMC-analysis of effective population sizes in molecular ecology and its application to black-and-white Ficedula flycatchers. Molecular Ecology 25:1058-1072.

303. Rönnegård, L., McFarlane, E.S., Husby, A., Kawakami, T., Ellegren, H. & Qvarnström, A. (2016) Increasing the power of genome wide association studies in natural populations using repeated measures - evaluation and implementation. Methods in Ecology and Evolution 7:792-799.

302. Bischof, R, Gregersen, E.R., Brøseth, H., Ellegren, H. & Flagstad, Ø. (2016) Non-invasive sampling reveals intra-sex territoriality in wolverines. Ecology and Evolution 6:1527-1536.

301. Kardos, M., Husby, A., McFarlane, S.E., Qvarnström, A. & Ellegren, H. (2016) Whole genome resequencing of extreme phenotypes in collared flycatchers highlights the difficulty of detecting quantitative trait loci in natural populations. Molecular Ecology Resources 16:216-227.

300. Bolivar, P., Mugal, C.F. Nater, A. & Ellegren, H. (2016) Recombination rate variation modulates gene sequence evolution mainly via GC-biased gene conversion, not Hill-Robertson interference, in an avian system. Molecular Biology and Evolution 33:216-227.


299. Mugal, C.F., Weber, C.C. & Ellegren, H. (2015) GC-biased gene conversion links the recombination landscape and demography to genomic base composition. BioEssays 37:1317-1326. 

298. Burri, R., Nater, A., Kawakami, T., Mugal, C.F., Olason, P.I., Smeds, L., Suh, A., Dutoit, L., Bureš, S., Garamszegi, L.Z., Hogner, S., Moreno, J., Qvarnström, A., Ružić, M., Sæther, S.A., Sætre, G.P., Török, J. & Ellegren, H. (2015) Linked selection and recombination rate variation drive the evolution of the genomic landscape of differentiation across the speciation continuum of Ficedula flycatchers. Genome Research 25:1656-1665.

297. Schmid, M., Smith, J., Burt, D.W., Aken, B.L., Antin, P.B., Archibald, A.L., Ashwell, C., Blackshear, P.J., Boschiero, C., Brown, C.T., Burgess, S.C., Cheng, H.H., Chow, W., Coble, D.J., Cooksey, A., Crooijmans, R.P.M.A., Damas, J., Davis, R.V.N., de Konig, D.-J., Delany, M.E., Derrien, T., Desta, T.T., Dunn, I.C., Dunn, M. Ellegren, H. et al. (79 additional co-authors). (2015) Third report on chicken genes and chromosomes. Cytogenetics and Genome Research 145:78-179.

296. Gervasi, V., Brøseth, H., Nilse, E.B., Ellegren, H., Flagstad, Ø. & Linnell, J.D.C. Compensatory immigration counteracts contrasting conservation strategies of wolverines (Gulo gulo) within Scandinavia. Biological Conservation 191:632-639.

295. Suh, A., Smeds, L. & Ellegren, H. (2015) The dynamics of incomplete lineage sorting across the ancient adaptive radiation of neoavian birds. PLoS Biology 13:e1002224.

294. Nater, A., Burri, R., Kawakami, T., Smeds, L. & Ellegren, H. (2015) Resolving evolutionary relationships in closely related species with whole-genome sequencing data. Systematic Biology 64:1000-1017.

293. Uebbing, S., Konzer, A., Xu, L., Backström, N., Brunström, B., Bergquist, J. & Ellegren, H. (2015) Quantitative mass spectrometry reveals partial translational regulation for dosage compensation in chicken. Molecular Biology and Evolution 32:2716-2725.

292. Knief, U., Schielzeth, H., Ellegren, H., Kempenaers, B. & Forstmeier, W. (2015) A prezygotic transmission distorter acting equally in female and male zebra finches Taeniopygia guttata. Molecular Ecology 24:3846-3859.

291. Smeds, L., Warmuth, V., Bolivar, P., Uebbing, S., Burri, R., Suh, A., Nater, A., Bureš, S., Garamszegi, L.Z., Hogner, S., Moreno, J., Qvarnström, A., Ružić, M., Sæther, S.A., Sætre, G.P., Török, J. & Ellegren H. (2015) Evolutionary analysis of the female-specific avian W chromosome. Nature Communications 6:7330.

290. Nadachowska-Brzyska, K., Li, C., Smeds, L., Zhang, G. & Ellegren, H. (2015) Temporal dynamics of avian populations during Pleistocene revealed by whole-genome sequences. Current Biology 25:1375-1380.

289. Husby, A., Kawakami, T., Rönnegård, L., Smeds, L., Ellegren, H. & Qvarnström, A. (2015) Genome-wide association mapping in a wild avian population identifies a link between genetic and phenotypic variation in a life-history trait. Proceedings of the Royal Society of London, Series B 282:20150156.

288. Mugal, C.D., Arndt, P.F, Holm, L. & Ellegren, H. (2015) Evolutionary consequences of DNA methylation on the GC content in vertebrate genomes. G3 5:441-447.

287. Jarvis, E.D., Mirarab, S., Aberer, A.J., Li, B., Houde, P., Li, C., Ho, S.Y., Faircloth, B.C., Nabholz, B., Howard, J.T., Suh, A., Weber, C.C., da Fonseca, R.R., Alfaro-Núñez, A., Narula, N., Liu, L., Burt, D., Ellegren, H., Edwards, S.V., Stamatakis, A., Mindell, D.P., Cracraft, J., Braun, E.L., Warnow, T., Jun, W., Gilbert, M.T. & Zhang G; Avian Phylogenomics Consortium. (2015) Phylogenomic analyses data of the avian phylogenomics project. Gigascience 4:4.


286. Jarvis, E.D. et al. (Avian Phylogenomics Consortiuum) (2014) Whole-genome analyses resolve early branches in the tree of life of modern birds. Science 346: 1320-1331.

285. Li, C. et al. (40 co-authors) (2014) Two Antarctic penguin genomes reveal insights into their evolutionary history and molecular changes related to the Antarctic environment. Gigascience 3:27.

284. Suh, A., Weber, C.C., Kehlmaier, C., Braun, E.L., Green, R.E., Fritz, U., Ray, D.A. & Ellegren, H. (2014) Crocodilian and turtle paleoviruses suggest Early Mesozoic coexistence of amniotes and Hepadnaviridae. PLoS Genetics, 10:e1004559.

283. Weber, C.C.,Boussau, B., Romiguier, J., Jarvis, E.D. & Ellegren, H. (2014) Evidence for GC-biased gene conversion as a driver of between-lineage differences in avian base composition. Genome Biology 15:549.

282. Weber, C.C., Nabholz, B., Romiguier, J. & Ellegren, H. (2014) Kr/Kc but not dN/dS correlates positively with body mass in birds, raising implications for inferring lineage-specific selection. Genome Biology 15:542.

281. Smeds, L., Kawakami, T., Burri, R., Bolivar, P., Husby, A., Qvarnström, A., Uebbing, S. & Ellegren, H. (2014) Genomic identification and characterization of the pseudoautosomal region in highly differentiated avian sex chromosomes. Nature Communications 5:5448.

280. Ekblom, R., Smeds, L. & Ellegren, H. (2014) Patterns of sequencing coverage bias revealed by ultra-deep sequencing of vertebrate mitochondria. BMC Genomics 15:467.

279. Kawakami, T., Smeds, L., Backström, N., Husby, A., Qvarnström, A., Mugal, C.F., Olason, P. & Ellegren, H. (2014) A high-density linkage map enables a second-generation collared flycatcher genome assembly and reveals the patterns of avian recombination rate variation and chromosomal evolution. Molecular Ecology 23:4035-4058.

278. Kawakami, T., Backström, N., Burri, R., Husby, A., Olason, P., Rice, A.M., Ålund, M., Qvarnström, A. & Ellegren, H. (2014) Estimation of linkage disequilibrium and interspecific gene flow in Ficedula flycatchers by a newly developed 50k SNP array. Molecular Ecology Resources 14:1248-1260.

277. Yazdi, H.P. & Ellegren, H. (2014) Old but not (so) degenerated: slow evolution of largely homomorphic sex chromosomes in ratites. Molecular Biology and Evolution 31:1444-1453.

276. Ellegren, H. (2014) Genome sequencing and population genomics in non-model organisms. Trends in Ecology and Evolution 29:51-63.


275. Nadachowska-Brzyska, K., Burri, R., Olason, P.I., Kawakami, T., Smeds, L. & Ellegren, H. (2013) Demographic divergence history of pied flycatcher and collared flycatcher inferred from whole-genome re-sequencing data. PLoS Genetics 9:e1003942

274. Uebbing, S., Künstner, A., Mäkinen, H. & Ellegren, H. (2013) Transcriptome sequencing reveals the character of incomplete dosage compensation across multiple tissues in flycatchers. Genome Biology and Evolution 5:1555-1566.

273. Ellegren, H. (2013) The evolutionary genomics of birds. Annual Reviews of Ecology, Evolution and Systematics 44:239-259.

272. Poelstra, J.W., Ellegren, H. & Wolf, J.B. (2013) An extensive candidate gene approach to speciation: diversity, divergence and linkage disequilibrium in candidate pigmentation genes across the European crow hybrid zone. Heredity 111:467-473.

271. Mugal, C.F., Arndt, P.F. & Ellegren, H. (2013) Twisted signatures of GC-biased gene conversion embedded in an evolutionary stable karyotype. Molecular Biology and Evolution 30:1700-1712.

270. Corl, A. & Ellegren, H. (2013) Sampling strategies for species trees: The effects on phylogenetic inference of the number of genes, number of individuals, and whether loci are mitochondrial, sex-linked, or autosomal. Molecular Phylogenetics and Evolution 67: 358-366.

269. Mugal, C.F., Nabholz, B. & Ellegren, H. (2013) Genome-wide analysis in chicken reveals that local levels of genetic diversity are mainly governed by the rate of recombination. BMC Genomics 14:86.

268. Adolfsson, S. & Ellegren, H. (2013) Lack of dosage compensation accompanies the arrested stage of sex chromosome evolution in ostriches. Molecular Biology and Evolution 30:806-810.

267. Backström, N., Saetre, G.P. & Ellegren, H. (2013) Inferring the demographic history of European Ficedula flycatcher populations. BMC Evolutionary Biology 13:2.

266. Parsch, J. & Ellegren, H. (2013) The evolutionary consequences of sex-biased gene expression. Nature Reviews Genetics 14:83-87.

265. Nabholz, B., Ellegren, H. & Wolf, J.B.W (2013) High levels of gene expression explain the strong evolutionary constraint of mitochondrial protein-coding genes. Molecular Biology and Evolution 30:272-284.


264. Ellegren, H., Smeds, L., Burri, R., Olason, P.I., Backström, N., Kawakami, T., Künstner, A., Mäkinen, H., Nadachowska-Brzysja, K., Qvarnström, A., Uebbing, S. & Wolf, J.B.W. (2012) The genomic landscape of species divergence in Ficedula flycatchers. Nature 491:756-760.

263. Knief, U., Schielzeth, H., Kempenaers, B., Ellegren, H. & Forstmeier, W. (2012) QTL and quantitative genetic analysis of beak morphology reveals patterns of standing genetic variation in the zebra finch. Molecular Ecology 21:3704-3717.

262. Nam, K. & Ellegren, H. (2012). Recombination drives vertebrate genome contraction. PLoS Genetics 8: e1002680.

261. Forstmeier, W., Schielzeth, H., Mueller, J.C., Ellegren, H. & Kempenaers, B. (2012) Heterozygosity-fitness correlations in zebra finches: microsatellite markers can be better than their reputation. Molecular Ecology 21: 3237-3249.

260. Corl, A. & Ellegren, H. (2012) The genomic signature of sexual selection in the genetic diversity of the sex chromosomes and autosomes. Evolution 66: 2138-2149.

259. Schielzeth, H., Forstmeier, W., Kempenaers, B. & Ellegren, H. (2012) QTL linkage mapping of wing length in zebra finch using genome-wide SNP markers. Molecular Ecology 21: 329-339.

258. Schielzeth, H., Kempenaers, B., Ellegren, H. & Forstmeier, W. (2012) QTL linkage mapping of zebra finch beak colour shows an oligogenic control of a sexually selected trait. Evolution 66: 18-30.


257. Künstner, A., Nabholz, B. & Ellegren, H. (2011) Significant selective constraint at 4-fold degenerate sites in the avian genome and its consequence for detection of positively selected genes. Genome Biology and Evolution 3: 1381-1389.

256. Ellegren, H. (2011) Emergence of male-biased genes on the chicken Z-chromosome: Contrasts between male and female heterogametic systems. Genome Research 21:2082-2086.

255. Mugal, C.F. & Ellegren, H. (2011) Substitution rate variation at human CpG sites correlates with non-CpG divergence, methylation level and GC content. Genome Biology 12:R58.

254. Künstner, A., Nabholz, B. & Ellegren, H. (2011). Evolutionary constraint in flanking regions of avian genes. Molecular Biology and Evolution 28:2481-2489.

253. Nabholz, B., Künstner, A., Wang, R., Jarvis, E.D. & Ellegren, H. (2011) Dynamic evolution of base composition: causes and consequences in avian phylogenomics. Molecular Biology and Evolution 28:2197-2210.

252. Ellegren, H. (2011) Sex chromosome evolution: recent progress and the influence of male and female heterogamety. Nature Reviews Genetics 12:157-166.

251. Rice, A.M., Rudh, A., Ellegren, H. & Qvarnström, A. (2011) A guide to the genomics of ecological speciation in natural animal populations. Ecological Letters 14:9-18.

250. Ellegren, H. (2011) The singing genome. Heredity 106:533-534.


249. Nabholz, B., Jarvis, E.D. & Ellegren, H. (2010) Obtaining mtDNA genomes from next-generation transcriptome sequencing: A case study of the basal Passerida (Aves: Passeriformes) phylogeny. Molecular Phylogenetics and Evolution 57: 466-470.

248. Backström, N., Lindell, J., Zhang, Y., Palkopoulou, E., Qvarnström, A., Saetre, G.P. & Ellegren, H. (2010) A high-density scan of the Z-chromosome in Ficedula flycatchers reveals candidate loci for diversifying selection. Evolution 64: 3461-3475.

247. Backström, N., Palkopoulou, E., Qvarnström, A. & Ellegren, H. (2010) No evidence for Z-chromosome rearrangements between the pied flycatcher and the collared flycatcher as judged by gene-based comparative genetic maps. Molecular Ecology 19: 3394-3405.

246. Axelsson, E., Albrechtsen, A., As P. van, Li, L., Megens, H.J., Vereijken, A.L.J., Crooijmans, R.P.M.A., Groenen, M.A.M., Ellegren, H., Willerslev, E. & Nielsen, R. (2010) Segregation distortion in chicken and the evolutionary consequences of female meiotic drive in birds. Heredity 105: 290-298.

245. Forstmeier, W. & Ellegren, H. (2010) Trisomy and triploidy are sources of embryo mortality in the zebra finch. Proceedings of the Royal Society of London, Series B 277: 2655-2660.

244. Nam, K., Mugal, C., Nabholz, B., Schielzeth, H., Wolf, J.B., Backström, N., Künstner, A., Balakrishnan, C.N., Heger, A., Ponting, C.P., Clayton, D.F. & Ellegren, H. (2010) Molecular evolution of genes in avian genomes. Genome Biology 11: R68.

243. Mank, J.E., Nam, K., Brunström, B. & Ellegren, H. (2010) Ontogenetic complexity of sexual dimorphism and sex-specific selection. Molecular Biology and Evolution 27: 1570-1578.

242. Qvarnström, A., Rice, A.M. & Ellegren, H. (2010) Speciation in Ficedula flycatchers. Philosophical Transactions of the Royal Society of London B: Biological Sciences 365: 1841-1852.

241. Vali, U., Dombrovski, V., Treinys, R., Bergmanis, U., Daroczi, S.J., Dravecky, M., Ivanovski, V., Lontkowski, J., Maciorowski, G., Meyburg, B.U., Mizera, T., Zeitz, R. & Ellegren, H. (2010) Widespread hybridization between the Greater Spotted Eagle Aquila clanga and the Lesser Spotted Eagle Aquila pomarina (Aves: Accipitriformes) in Europe. Biological Journal of the Linnean Society 100: 725-736.

240. Lira, J., Linderholm, A., Olaria, C., Brandström Durling, M., Gilbert, T.P., Ellegren, H., Willerslev, E., Lidén, K., Arsuaga, J.L. & Götherström, A. (2010) Ancient DNA reveals traces of Iberian Neolithic and Bronze Age lineages in modern Iberian horses. Molecular Ecology 19: 64-78.

239. Völker, M., Backström, N., Skinner, B.M., Langley, E.J., Bunzey, S.K., Ellegren, H. & Griffin, D.K. (2010) Copy number variation, chromosome rearrangement, and their association with recombination during avian evolution. Genome Research 20: 503-511.

238. Warren, W.C., Clayton, D.F., Ellegren, H., Arnold, A.P., Hillier, L.W., Künstner, A., Searle, S., White, S., Vilella, A.J., Fairley, S., Heger, A., Kong, L., Ponting, C.P., Jarvis, E.D., Mello, C.V., Minx, P., Lovell, P., Velho, T.A., Ferris, M., Balakrishnan, C.N., Sinha, S., Blatti, C., London, S.E., Li, Y., Lin, Y.C., George, J., Sweedler, J., Southey, B., Gunaratne, P., Watson, M., Nam, K., Backström, N., Smeds, L., Nabholz, B., Itoh, Y., Whitney, O., Pfenning, A.R., Howard, J., Völker, M., Skinner, B.M., Griffin, D.K., Ye, L., McLaren, W.M., Flicek, P., Quesada, V., Velasco, G., Lopez-Otin, C., Puente, X.S., Olender, T., Lancet, D., Smit, A.F., Hubley, R., Konkel, M.K., Walker, J.A., Batzer, M.A., Gu, W., Pollock, D.D., Chen, L., Cheng, Z., Eichler, E.E., Stapley, J., Slate, J., Ekblom, R., Birkhead, T., Burke, T., Burt, D., Scharff, C., Adam, I., Richard, H., Sultan, M., Soldatov, A., Lehrach, H., Edwards, S.V., Yang, S.P., Li, X., Graves, T., Fulton, L., Nelson, J., Chinwalla, A., Hou, S., Mardis, E.R. & Wilson, R.K. (2010) The genome of a songbird. Nature 464: 757-762.

237. Mugal, C.F., Wolf, J.B.W., von Grünberg H.H. & Ellegren, H. (2010) Conservation of neutral substitution rate and substitutional asymmetries in mammalian genes. Genome Biology and Evolution 2010: 19-28.

236. Künstner, A., Wolf, J.B.W., Backström, N., Whitney, O., Balakrishnan, C., Day, L., Edwards, S.V., Janes, D.E., Schlinger, B.A., Wilson, R.K., Jarvis, E.D., Warren, W.C. & Ellegren, H. (2010) Comparative genomics based on massive parallel transcriptome sequencing reveals patterns of substitution and selection across 10 bird species. Molecular Ecology 19 Suppl 1: 266-276.

235. Backström, N., Forstmeier, W., Schielzeth, H., Mellenius, H., Nam, K., Bolund, E., Webster, M.T., Öst, T., Schneider, M., Kempenaers, B. & Ellegren, H. (2010) The recombination landscape of the zebra finch Taeniopygia guttata genome. Genome Research 20: 485-495.

234. Ellegren, H. (2010) Evolutionary stasis – the stable chromosomes of birds. Trends in Ecology and Evolution 25: 283-291.

233. Mank, J.E., Nam, K. & Ellegren, H. (2010) Faster-Z evolution is predominantly due to genetic drift. Molecular Biology and Evolution 27: 661-670.

232. Brøseth, H., Flagstad, Ø., Wärdig, C., Johansson, M. & Ellegren, H. (2010) Large-scale noninvasive genetic monitoring of wolverines using scats reveals density dependent adult survival. Biological Conservation 143: 113-120.

231. Hansson, B., Ljungqvist, M., Dawson, D.A., Mueller, J.C., Olano-Marin, J., Ellegren, H. & Nilsson, J.Å. (2010) Avian genome evolution: insights from a linkage map of the blue tit (Cyanistes caeruleus). Heredity 104: 67-78.


230. Ellegren, H. (2009) Sex determination: two copies for one cock. Current Biology 19: R909-910.

229. Wolf, J.B.W., Künstner, A., Nam, K., Jakobsson, M. & Ellegren, H. (2009) Non-linear dynamics of non-synonymous (dN) and synonymous (dS) substitution rates affects inference of selection. Genome Biology and Evolution 1: 308-319.

228. Mank, J.E. & Ellegren, H. (2009) Sex-bias in gene expression is not the same as dosage compensation. Heredity 103: 434.

227. Ellegren, H. (2009) Is genetic diversity really higher in large populations? Journal of Biology 8: 41.

226. Ellegren, H. (2009) The different levels of genetic diversity in sex chromosomes and autosomes. Trends in Genetics 25: 278-284.

225. Hagenblad, J., Olsson, M., Parker, H.G., Ostrander, E.A. & Ellegren, H. (2009) Population genomics of the inbred Scandinavian wolf. Molecular Ecology 18: 1341-1351.

224. Mank, J.E. & Ellegren, H. (2009) Are sex-biased genes more dispensable? Biology Letters 23: 409-412.

223. Mank, J.E. & Ellegren, H. (2009) Sex-linkage of sexually antagonistic genes is predicted by female, but not male, effects in birds. Evolution 63: 1464-1472.

222. Axelsson, E. & Ellegren, H. (2009) Quantification of adaptive evolution of genes expressed in avian brain and the population size effect on the efficacy of selection. Molecular Biology and Evolution 26: 1073-1079.

221. Ellegren, H. (2009) A selection model of molecular evolution incorporating the effective population size. Evolution 63: 301-305.

220. Mank, J.E. & Ellegren, H. (2009) All dosage compensation is local: Gene-by-gene regulation of sex-biased gene expression on the chicken Z chromosome. Heredity 102: 312-320.

219. Ellegren, H. (2009) Genomic evidence for a large-Z effect. Proceedings of the Royal Society of London, Series B 276: 361-366.


218. Ellegren, H. (2008) Comparative genomics and the study of evolution by natural selection. Molecular Ecology 17: 4586-4596.

217. Berlin, S., Qu, L. & Ellegren, H. (2008) Adaptive evolution of gamete-recognition proteins in birds. Journal of Molecular Evolution 67: 488-496.

216. Brandström, M., Bradshaw, A.T., Gemmell, N.J. & Ellegren, H. (2008) The relationship between microsatellite polymorphism and recombination hotspots in the human genome. Molecular Biology and Evolution 25: 2579-2587.

215. Nam, K. & Ellegren, H. (2008) The chicken (Gallus gallus) Z chromosome contains at least three non-linear evolutionary strata. Genetics 180: 1131-1136.

214. Berlin, S., Qu, L. Yang, N. & Ellegren, H. (2008) Positive diversifying selection in avian Mx genes. Immunogenetics 60: 689-697.

213. Väli, U., Einarsson, A., Waitz, L. & Ellegren, H. (2008) To what extent do microsatellite markers reflect genome-wide genetic diversity in natural populations? Molecular Ecology 17: 3808-3817.

212. Ellegren, H. (2008) Platypus genome suggests a recent origin for the human X. Current Biology 18: R557-559.

211. Backström, N., Karaiskou, N., Leder, E.H., Gustafsson, L. Primmer, C.R., Qvarnström, A. & Ellegren, H. (2008) A gene-based genetic linkage map of the collared flycatcher (Ficedula albicollis) reveals extensive synteny and gene order conservation during 100 million years of avian evolution. Genetics 179: 1479-1495.

210. Brandström, M. & Ellegren, H. (2008) Genome-wide analysis of microsatellite polymorphism in chicken circumventing the ascertainment bias. Genome Research 18: 881-887.

209. Mank, J.E., Hultin-Rosenberg, L., Webster, M.T. & Ellegren, H. (2008) The unique genomic properties of sex-biased genes: insights from avian microarray data. BMC Genomics 9: 148.

208. Axelsson., E., Hultin-Rosenberg, L., Brandström, M., Zwahlén, M., Clayton, D.F. & Ellegren, H. (2008) Natural selection in avian protein-coding genes expressed in brain. Molecular Ecology 17: 3008-3017.

207. Ellegren, H. & Sheldon, B.C. (2008) Genetic basis for fitness differences in natural populations. Nature 452: 169-175.

206. Ellegren, H. (2008) Sequencing goes 454 and takes large-scale genomics into the wild. Molecular Ecology 17: 1629-1631.

205. Väli, U., Brandström, M., Johansson, M. & Ellegren, H. (2008) Insertion-deletion polymorphisms (indels) as genetic markers in natural populations. BMC Genetics 9: 8.

204. Hefti-Gautschi, B., Pfunder, M., Jenni, L. Keller, V. & Ellegren, H. (2008) Identification of conservation units in the European Merganser merganser based on nuclear and mitochondrial DNA markers. Conservation Genetics 10: 87-99.

203. Sundqvist, A.K., Ellegren, H. & Vilà, C. (2008) Wolf or dog? Genetic identification of predator from saliva collected around bite wounds on prey. Conservation Genetics 9: 1275-1279.

202. Backström, N., Fagerberg, S. & Ellegren, H. (2008) Genomics of wild bird populations: a gene-based set of references markers spread across the avian genome. Molecular Ecology 17: 964-980.

201. Mank, J.E., Hultin-Rosenberg, L., Zwahlén, M. & Ellegren, H. (2008) Pleiotropic constraint hampers the resolution of sexual antagonism in vertebrate gene expression. American Naturalist 171: 35-43.


200. Ellegren, H. & Parsch, J. (2007) The evolution of sex-biased genes and sex-biased gene expression. Nature Reviews Genetics 8: 689-698.

199. Mank, J.E., Hultin-Rosenberg, L., Axelsson, E. & Ellegren, H. (2007) Rapid evolution of female-biased, but not male-biased, genes expressed in the avian brain. Molecular Biology and Evolution 24: 2698-2706.

198. Ellegren, H., Hultin-Rosenberg, L., Brunström, B., Dencker, L., Kultima, K. & Scholtz, B. (2007) Faced with inequality: chicken does not have a general dosage compensation of sex-linked genes. BMC Biology 5: 40.

197. Brandström, M. & Ellegren, H. (2007) The genomic landscape of short insertion and deletion polymorphisms in the chicken (Gallus gallus) genome: a high frequency of deletions in tandem duplicates. Genetics 176: 1691-1701.

196. Mank. J.M. & Ellegren, H. (2007) Parallel divergence and degradation of the avian W sex chromosome. Trends in Ecology and Evolution 22: 389-391.

195. Ellegren, H. (2007) A dinosaur’s view on genome size evolution. Current Biology 17; R470-472.

194. Engen, S., Ringsby, T.H., Lande, R., Jensen, H., Sæther, B.E., Lillegard, M. & Ellegren, H. (2007) Effective size of fluctuating populations with overlapping generations: extension of Wright’s formula for two sexes. Evolution 61: 1873-1885.

193. Mank, J.M., Axelsson, E. & Ellegren, H. (2007) Fast-X on Z: rapid evolution of sex-linked genes in birds. Genome Research 17: 618-624.

192. Ellegren, H. (2007) Avian molecular evolutionary genomics. Cytogenetics and Genome Research 117: 120-130.

191. Ellegren, H. (2007) Characteristics, causes and evolutionary consequences of male-biased mutation. Proceedings of the Royal Society London, Series B 274: 1-10.

190. Kaiser, V.B., van Tuinen, M. & Ellegren, H. (2007) The phylogenetic position of the chicken revisited: insertion events of CR1 retrotransposable elements elucidate the order of speciation events in galliform species (Aves). Molecular Biology and Evolution 24: 338-347.

189. Hedmark, E. & Ellegren, H. (2007) DNA-based monitoring of two newly founded Scandinavian wolverine populations. Conservation Genetics 8: 843-852.

188. Hedmark, E., Persson, J. & Ellegren, H. (2007) Paternity and mating system in wolverines. Wildlife Biology 13: 13-30.


187. Kaiser, V.B. & Ellegren, H. (2006) Non-random distribution of genes with sex-biased expression in the chicken genome. Evolution 60: 1945-1951.

186. Backström, N., Brandström, M., Gustafsson, L., Qvarnström, A., Cheng, H. & Ellegren, H. (2006) Genetic mapping in a natural population of collared flycatchers (Ficedula albicollis): Conserved synteny but gene order rearrangements on the avian Z chromosome. Genetics 174: 377-386.

185. Backström, N., Qvarnström, A., Gustafsson, L. & Ellegren, H. (2006) Levels of linkage disequilibrium (LD) in a wild bird population. Biology Letters 2: 435-438.

184. Hailer, F., Helander, B., Folkestad, A.O., Ganusevich, S.A., Garstad, S., Hauff, P., Koren, C., Nygård, T., Volke, V., Vilà, C. & Ellegren, H. (2006) Bottlenecked but long-lived: high genetic diversity retained in white-tailed eagles upon recovery from population decline. Biology Letters 2: 316-319.

183. Webster, M.T., Axelsson, E. & Ellegren, H. (2006) Strong regional biases in nucleotide substitution in the chicken genome. Molecular Biology and Evolution 23: 1203-1216.

182. Sundqvist, A.K., Björnerfeldt, S., Leonard, J.A., Hailer, F., Hedhammar, Å., Ellegren, H. & Vilà, C. (2006) Unequal contribution of sexes in the origin of dog breeds. Genetics 172: 2345-2350.

181. Berlin, S., Brandström, M., Backström, N., Axelsson, E., Smith, N.G.C. & Ellegren, H. (2006) Substitution rate heterogeneity and the male mutation bias. Journal of Molecular Evolution 62: 226-233.

180. Berlin, S. & Ellegren, H. (2006) Accumulation of deleterious mutations on the female-specific W chromosome in birds. Journal of Molecular Evolution 62: 66-72.

179. Seddon, J.M., Sundqvist, A.K. & Ellegren, H. (2006) Genetic identification of immigrants in the Scandinavian wolf population. Conservation Genetics 7: 225-230.

178. Hedmark, E. & Ellegren, H. (2006) A test of the multiplex pre-amplification approach in microsatellite genotyping of wolverine faecal DNA. Conservation Genetics 7: 289-293.


177. Ellegren, H. (2005) The dog has its day. Nature 438: 745-746.

176. Ellegren, H. (2005) Natural selection in the evolution of human and chimps. Current Biology 15: R919-922.

175. Götherström, A., Anderung, C., Hellborg, L. Elburg, R., Smith, C. & Ellegren, H. (2005) Cattle domestication in the Near East was followed by hybridization with aurochs bulls in Europe. Proceedings of the Royal Society London, Series B 272: 2345-2350.

174. Hedmark, E. & Ellegren, H. (2005) Successful genotyping of DNA isolated from claws left on tanned carnivore hides. International Journal of Legal Medicine 119: 370-373.

173. Backström, N., Ceplitis, H. & Ellegren, H. (2005) Gene conversion drives the evolution of HINTW, a multi-copy gene family on the avian W chromosome. Molecular Biology and Evolution 22: 1992-1999.

172. Anderung, C., Bouwman, A., Persson, P., Carretero, J. M., Ortega, A. I., Arasuaga, J. L. Elburg, R., Smith, C. Ellegren, H. & Götherström, A. (2005) Genetics of Iberian Bronze Age cattle remains indicate prehistoric contacts over the Straits of Gibraltar. Proceedings of the National Academy of Sciences USA 102: 8431-8435.

171. Webster, M.T., Smith, N.G.C., Hultin-Rosenberg, L., Arndt, P.F. & Ellegren, H. Male-driven biased gene conversion governs the evolution of base composition in human Alu repeats. Molecular Biology and Evolution 22: 1468-1474.

170. Berggren, K.T., Ellegren, H., Hewitt, G.M. & Seddon, J.M. (2005) Understanding the phylogeographic patterns of European hedgehogs, Erinaceus concolor and E. europaeus using the MHC. Heredity 95: 84-90.

169. Ellegren, H. (2005) The avian genome uncovered. Trends in Ecology and Evolution 20: 180-186.

168. Vilà, C., Seddon, J.M. & Ellegren, H. (2005) Genes of domestic mammals augmented by backcrossing with wild ancestors. Trends in Genetics 21: 214-218.

167. Seddon, J. M., Parker, H.G., Ostrander, E.A. & Ellegren, H. (2005) SNPs in ecological and conservation studies: a test in the Scandinavian wolf population. Molecular Ecology 14: 503-511.

166. Axelsson, E., Webster, M.T., Smith, N.G., Burt, D.W. & Ellegren, H. (2005) Comparison of the chicken and turkey genomes reveals a higher rate of nucleotide divergence on microchromosomes than macrochromosomes. Genome Research 15: 120-125.


165. International Chicken Polymorphism Map Consortium. (2004) A genetic variation map for chicken with 2.8 million single nucleotide polymorphisms. Nature 432: 717-722.

164. International Chicken Genome Sequencing Consortium. (2004) Sequencing and comparative analysis of the chicken genome. Nature 432: 695-716.

163. Berlin, S. & Ellegren, H. (2004) Chicken W: a genetically uniform chromosome in a highly variable genome. Proceedings of the National Academy of Sciences USA 101: 15967-15969.

162. Seddon, J.M. & Ellegren, H. (2004) A temporal analysis shows MHC loci in the Scandinavian wolf population are consistent with neutral evolution. Proceedings of the Royal Society London, Series B 271: 2283-2291.

161. Smith, N.G., Brandström, M. & Ellegren, H. (2004) Evidence for turnover of functional noncoding DNA in mammalian genome evolution. Genomics 84: 806-813.

160. Webster, M.T., Smith, N.G.C., Lercher, M.J. & Ellegren, H. (2004) Gene expression, synteny, and local similarity in human noncoding mutation rates. Molecular Biology and Evolution 21: 1820-1830.

159. Saetre, P., Lindberg, J., Leonard, J., Olsson, K., Pettersson, U., Ellegren, H., Bergström, T., Vila, C. & Jazin, E. (2004) From wild wolf to domestic dog: gene expression changes in the brain. Brain Research Molecular Brain Research 126: 198-206.

158. Ellegren, H. (2004) Microsatellites: simple sequences with complex evolution. Nature Reviews Genetics 5: 435-445.

157. Sundström, H., Webster, M. T. & Ellegren, H. (2004) Reduced variation on the chicken Z chromosome. Genetics 167: 377-385.

156. Lindgren, G., Backström, N., Swinburne, J., Hellborg, L., Einarsson, A., Sandberg, K., Vilà, C., Binns, M. & Ellegren, H. (2004) Limited number of patrilines in horse domestication. Nature Genetics 36: 335-336.

155. Axelsson, E., Smith, N. G. C., Sundström, H., Berlin, S. & Ellegren, H. (2004) Male-biased mutation rate and divergence in autosomal, Z-linked and W-linked introns of chicken and turkey. Molecular Biology and Evolution 21: 1538-1547.

154. Jensen, H., Saether, B.E., Ringsby, T.H., Tufto, J., Griffith, S.C. & Ellegren, H. Lifetime reproductive success in relation to morphology in house sparrow (Passer domesticus). Journal of Animal Ecology 73: 599-611.

153. Hedmark, E., Flagstad Ø., Segerström P., Persson, J., Landa, A. & Ellegren, H. (2004) DNA-based individual and sex identification from wolverine (Gulo gulo) faeces and urine. Conservation Genetics 5: 405-410.

152. Hellborg, L., & Ellegren, H. (2004) Low levels of nucleotide diversity in mammalian Y chromosomes. Molecular Biology and Evolution 21: 158-163.

151. Flagstad, Ø., Hedmark, E., Landa, A., Brøseth, H., Persson, J., Andersen, R., Segerström, P. & Ellegren, H. (2004) Colonization history and non-invasive monitoring of a re-established wolverine (Gulo gulo) population. Conservation Biology 18: 676-688.

150. Ceplitis, H. & Ellegren, H. (2004) Adaptive molecular evolution in HINTW, a female-specific gene in birds. Molecular Biology and Evolution 21: 249-254.

149. Brohede, J., Arnheim, N. & Ellegren, H. (2004) Single molecule analysis of the hypermutable tetranucleotide repeat locus D21S1245 through sperm genotyping: a heterogeneous pattern of mutation but no clear male age effect. Molecular Biology and Evolution 21: 58-64.

148. Berlin, S., Smith, N.G.C. & Ellegren, H. (2004) Do avian mitochondria recombine? Journal of Molecular Evolution 58: 163-167.

147. Lawson-Handley, L., Ceplitis, H. & Ellegren, H. (2004) Evolutionary strata on the chicken Z chromosome: Implications to sex chromosome evolution. Genetics 167: 367-376.


146. Jensen, H., Saether, B.E., Ringsby, T.H., Tufto, J., Griffith, S.C. & Ellegren, H. (2003) Sexual variation in heritability and genetic correlations of morphological traits in house sparrow (Passer domesticus). Journal of Evolutionary Biology 16: 1296-1307.

145. Ellegren, H., Smith, N.G.C. & Webster, M.T. (2003) Mutation rate variation in the mammalian genome. Current Opinion in Genetics and Development 13: 562-568.

144. Brohede, J., Møller, A.P. & Ellegren, H. (2003) Individual variation in microsatellite mutation rate in barn swallows. Mutation Research 545: 73-80.

143. Rueness, E. K., Jorde, P. E., Hellborg, L., Stenseth, N. C., Ellegren, H. & Jacobsen, K. S. (2003) Cryptic population structure in a large, mobile mammalian predator: the Scandinavian lynx. Molecular Ecology 12: 2623-2633.

142. Rueness, E. K., Stenseth, N. C., O’Donoghue, M., Boutin, S., Ellegren, H. & Jakobsen, K. S. (2003) The phylogeography of the Canadian lynx: coinciding genetic and ecological spatial structuring. Nature 425: 69-72.

141. Bartosch-Härlid, A., Berlin, S., Smith, N.G.C., Møller, A.P. & Ellegren, H. (2003) Life history and the male mutation bias. Evolution 57: 2398-2406.

140. Ellegren, H. & Fridolfsson, A.K. (2003) Sex-specific mutation rates in salmonid fish. Journal of Molecular Evolution 56: 458-463.

139. Sundström, H., Webster, M.T. & Ellegren, H. (2003) Is the rate of insertion and deletion mutation male-biased? - Molecular evolutionary analysis of avian and primate sex chromosome sequences. Genetics 164: 259-268.

138. Flagstad, Ø., Walker, C.W., Vilà, C., Sundqvist, A.K., Fernholm, B., Hufthammar, A.K., Wiig, Ø., Kojola, I. & Ellegren, H. (2003) Two centuries of the Scandinavian wolf population: patterns of genetic variability and migration during an era of publication decline. Molecular Ecology 12: 869-880.

137. Ellegren, H. (2003) Levels of polymorphism on the sex-limited chromosome – a clue to Y from W? BioEssays 25: 163-167.

136. Vilà, C., Walker, C., Sundqvist, A.K., Flagstad, Ø., Andersone, Z., Casulli, A., Kojola, I., Valdmann, H., Halverson, J. & Ellegren, H. (2003) Combined use of maternal, paternal and bi-parental genetic markers for the identification of wolf-dog hybrids. Heredity 90: 17-24.

135. Smith, N.G.C., Webster, M.T. & Ellegren, H. (2003) A low rate of simultaneous double nucleotide mutations in primates. Molecular Biology and Evolution 20: 47-53.

134. Webster, M.T., Smith, N.G.C. & Ellegren, H. (2003) Compositional evolution of non-coding DNA in the primate genome. Molecular Biology and Evolution 20: 278-286.

133. Vila, C., Sundqvist, A.K., Flagstad, Ø., Seddon, J.M., Bjornerfeldt, S., Kojola, I., Sand, H., Wabakken, P. & Ellegren, H. (2003) Rescue of a severely bottlenecked wolf (Canis lupus) population by a single immigrant. Proceedings of the Royal Society of London Series B 270: 91-98.

132. Hellborg, L. & Ellegren, H. (2003) Y chromosome conserved anchored tagged sites (YCATS) for the analysis of mammalian male-specific sequences. Molecular Ecology 12: 283-291.


131. Hurst, L.D. & Ellegren, H. (2002) Mystery of the mutagenic male. Nature 420: 365-366.

130. Ellegren, H. (2002) Mismatch repair and mutation biases in microsatellite DNA. Trends in Genetics 18: 552.

129. Seddon, J.M. & Ellegren, H. (2002) MHC class II genes in European wolves: a comparison with dogs. Immunogenetics 54: 590-600.

128. Hipkiss, T., Hörnfeldt, B., Lundmark, A., Norbäck, M. & Ellegren, H. (2002) Sex ratio and age structure of nomadic Tengmalm’s owls: a molecular approach. Journal of Avian Biology 33: 107-110.

127. Ellegren, H. (2002) It took many mares to form the domestic horse. Trends in Genetics 18:500-501.

126. Smith, N.G.C., Webster, M.T. & Ellegren, H. (2002) Deterministic mutation rate variation in the human genome. Genome Research 12: 1350-1356.

125. Webster, M., Smith, N.G.C. & Ellegren, H. (2002) Microsatellite evolution inferred from human-chimpanzee genomic sequence alignment at the Mb-scale level. Proceedings of the National Academy of Sciences USA 99: 8748-8753.

124. Ellegren, H. (2002) Human mutation – blame (mostly) men. Nature Genetics 31: 9-10.

123. Brohede, J., Primmer C.R., Møller A.P. & Ellegren H. (2002) The rate and pattern of germline mutation differ between individual microsatellite loci. Nucleic Acids Research 30: 1997-2003.

122. Ellegren, H. (2002) Microsatellite evolution: a battle between replication slippage and point mutations. Trends in Genetics 18: 70.

121. Ellegren, H. (2002) Dosage compensation: do birds do it as well? Trends in Genetics 18: 25-28.

120. Hellborg, L., Walker, C.W., Rueness, E.K., Stacy, J.E., Kojola, I., Valdmann, H., Vilá, C., Zimmermann, B., Jakobsen, K.S. & Ellegren, H. (2002) Differentiation and levels of genetic variation in northern European lynx (Lynx lynx) populations revealed by microsatellites and mitochondrial DNA analysis. Conservation Genetics 3: 97-111.


119. Montell, H., Fridolfsson, A.K. & Ellegren, H. (2001) Contrasting levels of nucleotide diversity on the avian Z and W sex chromosomes. Molecular Biology and Evolution 18: 2010-2016.

118. Berlin, S. & Ellegren, H. (2001) Clonal co-transmission of mitochondrial DNA and the female-specific avian W chromosome. Nature 413: 37-38.

117. Sundqvist, A.K., Ellegren, H., Olivier, M. & Vilá, C. (2001) Y chromosome haplotyping in Scandinavian wolves (Canis lupus) based on microsatellite markers. Molecular Ecology 10: 1959-1966.

116. Hedrick, P., Fredrickson, R. & Ellegren, H. (2001) Evaluation of d2, a microsatellite measure of inbreeding and outbreeding, in wolves with a known pedigree. Evolution 6:1256-1260.

115. Lindgren, G, Swinburne, J.E., Breen, M., Mariat, D., Sandberg, K., Guerin, G., Ellegren, H., & Binns, M.M. (2001) Physical anchorage and orientation of equine linkage groups by FISH mapping BAC clones containing microsatellite markers. Animal Genetics 32: 37-39.

114. Lindgren, G., Breen, M., Godard, S., Bowling, A., Murray, J., Scavone, M., Skow, L., Sandberg, K., Guerin, G., Binns, M. & Ellegren, H. (2001) Mapping of 13 horse genes by fluorescence in-situ hybridization (FISH) and somatic cell hybrid analysis. Chromosome Research 9: 53-59.

113. Ellegren, H. (2001) Hens, cocks and avian sex determination - A quest for genes on Z or W? EMBO Reports 2: 192-196.

112. Vila, C., Ellegren, H,, Götherström, A., Leonard, J.A. & Wayne, R.K. (2001) Tales from the DNA of domestic horses – Reply. Science 292: 218-219.

111. Vilà, C., Leonard, J.A., Götherström, A., Marklund, S., Sandberg, K., Lidén, K., Wayne, R.K. & Ellegren, H. (2001) Widespread origins of domestic horse lineages. Science 291: 474-477.

110. Saetre, G.P., Borge, T., Lindell, J., Moum, T., Primmer, C.R., Sheldon, B.C., Haavie, J., Johnsen, A. & Ellegren. H. (2001) Speciation, introgressive hybridization and nonlinear rate of molecular evolution in flycatchers. Molecular Ecology 10: 737-749.

109. Ellegren, H. & Carmichael, A. (2001) Multiple and independent cessation of recombination between avian sex chromosomes. Genetics 158: 325-331.

108. Walker, C.W., Vilà, C., Landa, A., Lindén, M. & Ellegren, H. (2001) Genetic variation and structure in Scandinavian wolverine (Gulo gulo) populations. Molecular Ecology 10: 53-62.


107. Ellegren, H. (2000) Microsatellite mutations in the germline: implications for evolutionary inference. Trends in Genetics 16: 551-558.

106. Carmichael, A.N., Fridolfsson, A.K. & Ellegren, H. (2000) Male-biased mutation rates revealed from Z- and W-chromosome linked ATP synthase a-subunit (ATP5A1) sequences in birds. Journal of Molecular Evolution 50: 443-447.

105. Fridolfsson, A.K. & Ellegren, H. (2000) Molecular evolution of the avian CHD1 genes on the Z and W sex chromosomes. Genetics 155: 1903-1912.

104. Hörnfeldt, B, Hipkiss, T, Fridolfsson, AK, Eklund, U & Ellegren, H. (2000) Sex ratio and fledging success of supplementary-fed Tengmalm's owl broods. Molecular Ecology 9: 187-192.

103. Ellegren, H. (2000) Heterogeneous mutation processes in human microsatellite DNA sequences. Nature Genetics 24: 400-402.

102. Ellegren, H. (2000) Evolution of the avian sex chromosomes and their role in sex determination. Trends in Ecology and Evolution 15: 188-192.

101. Berlin, S., Merila, J. & Ellegren, H. (2000) Isolation and characterization of polymorphic microsatellite loci in the common frog, Rana temporaria. Molecular Ecology 9: 1938-1939.


100. Vila, C., Amorim, I.R., Leonard, J.A., Posada, D., Castroviejo, J., Petrucci-Fonseca, F., Crandall, K.A., Ellegren, H. & Wayne, R.K. (1999) Mitochondrial DNA phylogeography and population history of the grey wolf Canis lupus. Molecular Ecology 8: 2089-2103.

99. Guérin, G. et al. (33 additional authors) (1999) Report of the international equine gene mapping work shop: male linkage map. Animal Genetics 30: 341-354. 98. Ellegren, H. (1999) Inbreeding and relatedness in Scandinavian grey wolves. Hereditas 130: 239-244.

97. Ellegren, H., Carlsson, A. & Stenberg, I. (1999) Genetic structure and variability of white-backed woodpecker (Dendrocopus leucotos) populations in northern Europe. Hereditas 130: 291-299.

96. Saino, N., Ellegren, H. & Møller, A.P. (1999) No evidence for adjustment of sex allocation in relation to paternal ornamentation and paternity in barn swallows. Molecular Ecology 8: 399-406.

95. Brohede, J. & Ellegren, H. (1999) Microsatellite evolution: polarity of substitutions within repeats and neutrality of flanking sequences. Proceedings of the Royal Society London, Series B 266: 825-833.

94. Saino, N., Primmer, C., Ellegren, H. & Møller, A.P. (1999) Breeding asynchrony affects certainty of paternity in the barn swallow (Hirundo rustica). Behavioural Ecology and Sociobiology 45: 211-218.

93. Fridolfsson, A.K. & Ellegren, H. (1999) A simple and universal method for sexing non-ratite birds. Journal of Avian Biology 30: 116-121.

92. Lindgren, G., Persson, H. & Ellegren, H. (1999) Five equine dinucleotide microsatellite loci HTG17, HTG20, HTG21, HTG28 and HTG31. Animal Genetics 30: 70-71.


91. Johnsen, A., Lifjeld, J.T., Rohde, P.A. & Ellegren, H. (1998) Sexual conflict over fertilizations: female bluethroats escape male paternity guards. Behavioural Ecology and Sociobiology 43: 401-408.

90. Sheldon, B.C. & Ellegren, H. (1998) Sexual selection resulting from extra-pair paternity in collared flycatchers. Animal Behaviour 57:285-298.

89. Lidén, K., Götherström, A. & Ellegren, H. (1998) "Alia vero gens ibi moratur Suehans, quae velud Thyringi equis utuntur eximiis" or the excellent horses in Sveland. Laborativ Arkeologi 10-11: 59-64.

88. Hurst, L.D. & Ellegren, H. (1998) Sex differences in the mutation rate. Trends in Genetics 14: 446-452.

87. Sheldon, B.C. & Ellegren, H. (1998) Paternal effort related to experimentally manipulated paternity of male collared flycatchers. Proceedings of the Royal Society London, Series B 265: 1737-1742.

86. Sheldon, B.C., Merilä, J., Lindgren, G. & Ellegren, H. (1998) Gender and environmental sensitivity in nestling collared flycatchers. Ecology 79: 1939-1948.

85. Knott, S.A., Marklund, L., Haley, C.S., Andersson, K., Davies, W., Ellegren, H., Fredholm, M., Hoyheim, B., Hansson, I., Lundström, K., Möller, M. & Andersson, L. (1998) Multiple marker mapping of quantitative trait loci in an outbred cross between wild boar and large white pigs. Genetics 149: 1069-1080.

84. Lindgren, G., Sandberg, K., Persson, H., Marklund, S., Breen, M., Sandgren, B., Carlstén, J. & Ellegren, H. (1998) A primary male autosomal linkage map of the horse genome. Genome Research 8: 951-966.

83. Fridolfsson, A.K., Cheng, H., Copeland, N.G., Jenkins, N.A., Liu, H.C., Raudsepp, T., Woodage, T., Chowdhary, B., Halverson, J. & Ellegren, H. (1998) Evolution of the avian sex chromosomes from an ancestral pair of autosomes. Proceedings of the National Academy of Sciences USA 95: 8147-8152.

82. Merilä, J., Sheldon, B.C. & Ellegren, H. (1998) Quantitative genetics of sexual dimorphism in the collared flycatcher Ficedula albicollis. Evolution 52: 870-876.

81. Primmer, C.R., Saino, N., Møller, A.P. & Ellegren, H. (1998) Unravelling the process of microsatellite evolution through analysis of germline mutations in barn swallows. Molecular Biology and Evolution 15: 1047-1054.

80. Primmer, C.R. & Ellegren, H. (1998) Patterns of molecular evolution at avian microsatellite loci. Molecular Biology and Evolution 15: 997-1008.

79. Lifjeld, J.T., Slagsvold, T. & Ellegren, H. (1998) Experimentally reduced paternity affects paternal effort and reproductive success in pied flycatchers. Animal Behaviour 55: 319-329.


78. Ellegren, H. & Fridolfsson, A.K. (1997) Male-driven evolution of DNA sequences in birds. Nature Genetics 17:182-184.

77. Ellegren, H., Lindgren, G., Primmer, C.R. & Møller, A.P. (1997) Fitness loss and germline mutations in barn swallows breeding in Chernobyl. Nature 389: 393-396.

76. Hasler-Rapacz, J.H., Ellegren, H., Fridolfsson, A.K., Kirkpatrick, B., Kirk, S., Andersson, L. & Rapacz, J. (1997) Positional candidate cloning of a gene for familial hypercholesterolemia in swine. American Journal of Medical Genetics 76:379-386.

75. Fridolfsson, A.K., Hori, T., Winterø, A.K., Fredholm, M., Yerle, M., Robic, A., Andersson, L. & Ellegren, H. (1997) Expansion of the pig comparative map by expressed sequenced tags (EST) mapping. Mammalian Genome 8: 907-912.

74. Merilä, J., Sheldon, B.C. & Ellegren, H. (1997) Antagonistic natural selection revealed by molecular sex identification of nestling collared flycatchers. Molecular Ecology 6: 1167-1175.

73. Ellegren H. & Sheldon B.C. (1997) Flow cytometry for sexing birds – Reply. Trends in Ecology and Evolution 12: 489-490.

72. Ellegren, H., Moore, S., Robinson, N., Byrne, K., Ward, W. & Sheldon, B.C. (1997) Microsatellite evolution - a reciprocal study of repeat lengths at homologous loci in cattle and sheep. Molecular Biology and Evolution 14: 854-860.

71. Primmer, C.R., Raudsepp, T., Chowdhary, B. & Ellegren, H. (1997) Low frequency of microsatellites in the avian genome. Genome Research 7: 471-482.

70. Ellegren, H. & Sheldon, B. (1997) New tools for sex identification and the study of sex allocation in birds. Trends in Ecology and Evolution 12: 255-259.

69. Breen, M., Lindgren, G., Binns, M.M., Norman, J., Irvin, Z., Bell, K., Sandberg, K. & Ellegren, H. (1997) Genetical and physical assignments of equine microsatellite - first integration of anchored markers in horse genome mapping. Mammalian Genome 8: 267-273.

68. Saino, N., Primmer, C.R., Ellegren, H. & Møller, A.P. (1997) An experimental study of paternity and tail ornamentation in the barn swallow. Evolution 51: 562-570.

67. Lifjeld, J.T., Slagsvold, T. & Ellegren, H. (1997) Experimental mate switching in pied flycatchers: male copulatory access and fertilization success. Animal Behaviour 53: 1225-1232.

66. Lifjeld, J.T., Slagsvold, T., Dale, S. & Ellegren, H. (1997) A sexually selected paradox in the pied flycatcher: attractive males are cuckolded. Auk 114: 112-115.

65. Sheldon, B.C., Merilä, J., Qvarnström, A. & Ellegren, H. (1997) Female genetic benefit from extra-pair copulation predicted by relative size of male secondary sexual character. Proceedings of the Royal Society London, Series B 264: 297-302.


64. Ellegren, H. (1996) First gene on the avian W chromosome provides a tag for universal sexing of non-ratite birds. Proceedings of the Royal Society London, Series B 263: 1635-1641.

63. Ellegren, H., Savolainen, P. & Rosén, B. (1996) The genetical history of an isolated population of the endangered grey wolf Canis lupus: a study using nuclear and mitochondrial polymorphisms. Philosophical Transactions of the Royal Society London, Series B 351: 1661-1669.

62. Sheldon, B.C. & Ellegren, H. (1996) Offspring sex ratio and paternity in collared flycatcher. Proceedings of the Royal Society London, Series B 263: 1017-1021.

61. Primmer, C.R., Møller, A.P. & Ellegren, H. (1996) New microsatellites from the pied flycatcher and the swallow genomes. Hereditas 124: 281-283.

60. Ellegren, H., Gustafsson, L. & Sheldon, B.C. (1996) Sex ratio adjustment in relation to paternal attractiveness in a wild bird population. Proceedings of the National Academy of Sciences USA 93: 11723-11728.

59. Primmer, C.R., Saino, N., Møller, A.P. & Ellegren, H. (1996) Directional evolution in germline microsatellite mutations. Nature Genetics 13: 391-393.

58. Marklund, L., Johansson Möller, M., Høyheim, B., Davies, W., Fredholm, M., Juneja, R.K., Mariani, P., Coppieters, W., Ellegren, H. & Andersson, L. (1996) A comprehensive linkage map of the pig based on a wild pig - Large White intercross. Animal Genetics 27: 255-269.

57. Mariani, P., Johansson Möller, M., Høyheim, B., Marklund, L., Davies, W., Ellegren, H. & Andersson, L. (1996) Assignment of the extension (E) coat color locus to pig chromosome 6. Journal of Heredity 87: 272-276.

56. Primmer, C., Møller, A.P., & Ellegren, H. (1996) A wide-range survey of microsatellite cross-species amplification in birds. Molecular Ecology 5: 365-378.

55. Ellegren, H., Mikko, S., Wallin, K. & Andersson, L. (1996) Limited polymorphism at major histocompatibility complex (MHC) loci in the Swedish moose. Molecular Ecology 5: 3-10.


54. Larsen, N.J., Ellegren, H., Bräuner-Nielsen, P. & Andersson, L. (1995) Genetic variation at the growth hormone locus in a wild pig intercross; test of association to phenotypic traits and linkage to the blood group D locus. Theoretical and Applied Genetics 91: 1074-1077.

53. Ellegren, H., Primmer, C.R. & Sheldon, B.C. (1995) Microsatellite evolution: directionality or bias? Nature Genetics 11: 360-362.

52. Ellegren, H., Lifjeld, J.T., Slagsvold, T. & Primmer, C.R. (1995) Handicapped males and extrapair paternity in pied flycatchers: a study using microsatellite markers. Molecular Ecology 4: 441-447.

51. Paszek, A.A. et al. (a total of 21 authors) (1995) First international workshop on porcine chromosome 6. Animal Genetics 26: 377-401.

50. Ellegren, H. & Basu, T. (1995) Filling the gaps in the porcine linkage map: isolation of microsatellites from chromosome 18 using flow-sorting and SINE-PCR. Cytogenetics and Cell Genetics 71: 370-373.

49. Edfors-Lilja, I., Gustafsson, U., Duval-Iflah, Y., Ellegren, H., Johansson, M., Juneja, R.K., Marklund, L. & Andersson, L. (1995) The porcine intestinal receptor for Escherichia coli K88ab, K88ac; Regional localization on chromosome 13 and influence in IgG response to the K88 antigen. Animal Genetics 26: 237-242.

48. Primmer, C., Møller, A.P., & Ellegren, H. (1995) Resolving genetic relationships with microsatellite markers: A parentage testing system for the swallow Hirundo rustica. Molecular Ecology 4: 493-498.

47. Ellegren, H. (1995) Mutation rates at porcine microsatellite loci. Mammalian Genome 6: 376-377.

46. Archibald, A. et al. (a total of 50 authors). (1995) The PiGMaP consortium linkage map of the pig (Sus scrofa). Mammalian Genome 6: 157-175.

45. Johansson, M., Ellegren, H. & Andersson, L. (1995) Comparative mapping reveals extensive linkage conservation but also gene order rearrangements between the pig and human genomes. Genomics 25: 682-690.


44. Johansson, M., Ellegren, H., Marklund, L., Coppieters, W. & Andersson, L. (1994) Linkage maps of porcine chromosomes 3, 6 and 9 in pigs based on 31 polymorphic markers. Mammalian Genome 5: 785-790.

43. Ellegren, H., Chowdhary, B.P., Fredholm, M., Høyheim, B., Johansson, M., Bräuner-Nielsen, P., Thomsen, P.D. & Andersson, L. (1994) A physically anchored linkage map of pig chromosome 1 uncovers sex- and position-specific recombination rates. Genomics 24: 342-350.

42. Andersson, L., Edfors-Lilja, I., Ellegren, H., Johansson, M. & Marklund, L. (1994) Mapping trait loci by crossbreeding genetically divergent populations of domestic animals. Animal Biotechnology 5: 225-231.

41. Ellegren, H., Chowdhary, B.P., Johansson, M., Marklund, L., Fredholm, M., Gustavsson, I. & Andersson, L. (1994) A primary linkage map of the porcine genome reveals a low rate of genetic recombination in the pig. Genetics 137: 1089-1100.

40. Ellegren, H., Chowdhary, B., Johansson, M. & Andersson, L. (1994) Integrating the porcine physical and linkage map using cosmid-derived markers. Animal Genetics 25: 155-164.

39. Ellegren, H., Johansson, M., Hartman, G. & Andersson, L. (1994) DNA fingerprinting with the human 33.6 minisatellite probe identifies sex in beavers Castor fiber. Molecular Ecology 3: 273-274.

38. Andersson, L., Haley, C.S., Ellegren, H., Knott, S.A., Johansson, M., Andersson, K., Andersson-Eklund, L., Edfors-Lilja, I., Fredholm, M., Hansson, I., Håkansson, J. & Lundström, K. (1994) Genetic mapping of quantitative trait loci for growth and fatness in pigs. Science 263: 1771-1774.

37. Chowdhary, B.P., Ellegren, H., Johansson, M., Andersson, L. & Gustavsson, I. (1994) In situ hybridization mapping of the growth hormone receptor (GHR) gene assigns a linkage group (C9, FSA, GHR and S0105) to chromosome 16 in pigs. Mammalian Genome 5: 160-162.

36. Ellegren, H. (1994) Linkage mapping of the APOA1 gene to pig chromosome 9. Mammalian Genome 5: 58-59.

35. Marklund, S., Ellegren, H., Eriksson, S., Sandberg, K. & Andersson, L. (1994) Parentage testing and linkage analysis in the horse using a set of highly polymorphic microsatellites. Animal Genetics 25: 19-23.


34. Ellegren, H. (1993) Speed of migration and migratory flight lengths of passerine birds ringed during autumn migration in Sweden. Ornis Scandinavica 24: 220-228.

33. Holmgren, N., Ellegren, H. & Pettersson, J. (1993) Stopover length, body mass and fuel deposition rate in autumn migrating adult Dunlins Calidris alpine: evaluating the effects of moulting status and age. Ardea 81: 9-20.

32. Holmgren, N., Ellegren, H. & Pettersson, J. (1993) The adaptation of moult pattern in migratory Dunlins Calidris alpine. Ornis Scandinavica 24: 21-27.

31. Ellegren, H. (1993) Abundant (A)n•(T)n mononucleotide repeats in the pig genome: linkage mapping of the porcine APOB, FSA, ALOX12, PEPN and RLN loci. Animal Genetics 24: 367-372.

30. Edfors-Lilja, I., Ellegren, H., Winterø, A.K., Ruohonen-Lehto, M., Fredholm, M., Gustavsson, U., Juneja, R.K. & Andersson, L. (1993) A large linkage group on pig chromosome 7 including the MHC class I, class II (DQB), and class III (TNFB) genes. Immunogenetics 38: 363-366.

29. Ellegren, H., Hartmann, G., Johansson, M. & Andersson, L. (1993) Major histocompatibility complex monomorphism and low levels of DNA fingerprinting variability in a reintroduced and rapidly expanding population of beavers. Proceedings of the National Academy of Sciences (USA) 90: 8150-8153.

28. Chowdhary, B.P., Johansson, M., Chaudhary, R., Ellegren, H., Gu, F., Andersson, L. & Gustavsson, I. (1993) In situ hybridization mapping and RFLP analysis of the porcine albumin (Alb) and transferrin (Tf) genes. Animal Genetics 24: 85-90.

27. Johansson, M., Chowdhary, B., Gu, F., Ellegren, H., Gustavsson, I. & Andersson, L. (1993) Genetic analysis of the gene for porcine submaxillary gland mucin (MUC): physical assignment of the MUC and interferon gamma (IFNG) genes to chromosome 5. Journal of Heredity 84: 259-262.

25. Ellegren, H. (1993) Variable SINE 3' poly(A) sequences in the pig genome: an abundant class of genetic markers. Mammalian Genome 4: 429-434.

26. Ellegren, H. (1993) Genomic DNA from museum bird feathers. In: Ancient DNA (B. Herrman & S. Hummel, Eds.), pp. 207-213, Springer Verlag, New York.

24. Ellegren, H., Fredholm, M., Edfors-Lilja, I., Winterø, A.K. & Andersson, L. (1993) Conserved synteny between pig chromosome 8 and human chromosome 4 but rearranged and distorted linkage maps. Genomics 17: 599-603.

23. Ellegren, H., Davies, C.J. & Andersson, L. (1993) Strong association between polymorphisms in an intronic microsatellite and in the coding sequence of the BoLA-DRB3 gene. Implications for microsatellite stability and PCR-based DRB3 typing. Animal Genetics 24: 269-276.

22. Ellegren, H., Johansson, M., Chowdhary, B.P., Marklund, S., Brunnberg, L., Bräuner-Nielsen, P., Edfors-Lilja, I., Gustavsson, I., Juneja, R.K. & Andersson, L. (1993) Assignment of 20 microsatellite markers to the porcine linkage map. Genomics 16: 431-439.

21. Lagercrantz, U., Ellegren, H. & Andersson, L. (1993) The abundance of various polymorphic simple repeat motifs differ between plants and vertebrates. Nucleic Acids Research 21: 1111-1115.


20. Ellegren, H. (1992) Effects of age and sex on the fat-free body mass in autumn migrating Bluethroats Luscinia s. svecica. Ardea 80: 255-259.

19. Ellegren, H. & Fransson, T. (1992) Fat loads and estimated flight ranges in four Sylvia warblers analysed on autumn migration at Gotland, SE Sweden. Ringing & Migration 13: 1-12.

18. Johansson, M., Ellegren, H., Marklund, L., Gustavsson, U., Ringmar-Cederberg, E., Andersson, K., Edfors-Lilja, I. & Andersson, L. (1992) The gene for dominant white color in the pig is closely linked to ALB and PDGFRA on chromosome 8. Genomics 14: 965-969.

17. Ellegren, H. (1992) Polymerase chain reaction (PCR) analysis of avian microsatellites - a new approach to genetic studies in birds. Auk 109: 886-895.

16. Ellegren, H. (1992) Population genetics of museum birds by hypervariable microsatellite typing of genomic DNA prepared from museum feathers. Ancient DNA Newsletter 1: 10.

15. Johansson, M., Ellegren, H. & Andersson, L. (1992) Cloning of porcine microsatellites. Journal of Heredity 83: 196-198.

14. Ellegren, H., Johansson, M., Sandberg, K. & Andersson, L. (1992) Cloning of highly polymorphic microsatellites in the horse. Animal Genetics 23: 133-142.

13. Ellegren, H., Andersson, L., Johansson, M. & Sandberg, K. (1992) DNA fingerprinting in horses using a simple (TG)n probe and its application to population comparisons. Animal Genetics 23: 1-9.

12. Mariani, P., Johansson, M., Ellegren, H., Harbitz, I., Juneja, R.K. & Andersson, L. (1992) Multiple RFLPs in the porcine calcium release channel gene (CRC): assignment to the halothane (HAL) linkage group. Animal Genetics 23: 257-262.


11. Ellegren, H. (1991) DNA typing of museum birds. Nature 354: 113.

10. Ellegren, H. (1991) Stopover ecology in autumn migrating Bluethroats Luscinia s. svecica in relation to age and sex. Ornis Scandinavica 22: 340-348.

9. Ellegren, H. & Wallin, K. (1991) Autumn migrating Bluethroats Luscinia s. svecica orient in a east-southeastern direction at Gävle, East Sweden. Ornis Svecica 1: 47-50.

8. Ellegren, H., Andersson, L. & Wallin, K. (1991) DNA polymorphism in the moose (A. alces) revealed by the polynucleotide probe (TC)n. Journal of Heredity 82: 429-431.

7. Ellegren, H. (1991) Fingerprinting birds DNA with a polynucleotide probe (TG)n. Auk 108: 956-958.

6. Mariani, P., Johansson, M., Ellegren, H. & Andersson, L. (1991) A TaqI RFLP at the porcine platelet-derived growth factor ß-receptor locus (PDGFRB). Animal Genetics 22: 446.

5. Mariani, P., Johansson, M., Ellegren, H. & Andersson, L. (1991) TaqI and PvuII RFLPs at the porcine plasminogen activator, urokinase, locus (PLAU). Animal Genetics 22: 447.

4. Kågedahl, L., Engström, B., Ellegren, H., Lieber, A.K., Lundström, H., Persson, A. & Schenning, M. (1991) Chemical, physical and chromatographic properties of Superdex 75 prep grade and Superdex 200 prep grade gel filtration media. Journal of Chromatography 537: 17-32.


3. Ellegren, H. (1990) Timing of autumn migration in Bluethroats Luscinia s. svecica depends on timing of breeding. Ornis Fennica 67: 13-17.

2. Ellegren, H. (1990) Autumn migration speed in Scandinavian Bluethroats Luscinia s. svecica. Ringing & Migration 11: 121-131.


1. Ellegren, H. & Låås, T. (1989) Size-exclusion chromatography of DNA restriction fragments; fragment length determinations and a comparison with the behaviour of proteins in size-exclusion chromatography. Journal of Chromatography 467: 217-226.